Burkholderia species are a vast group of human pathogenic, phytopathogenic, and plant- or environment-associated bacteria. B. pseudomallei, B. mallei, and B. cepacia complex are the causative agents of melioidosis, glanders, and cystic fibrosis-related infections, respectively, which are fatal diseases in humans and animals. Due to their high resistance to antibiotics, high mortality rates, and increased infectivity via the respiratory tract, B. pseudomallei and B. mallei have been listed as potential bioterrorism agents by the Centers for Disease Control and Prevention. Burkholderia species are able to produce a large network of surface-exposed polysaccharides, i.e., lipopolysaccharides, capsular polysaccharides, and exopolysaccharides, which are virulence factors, immunomodulators, major biofilm components, and protective antigens, and have crucial implications in the pathogenicity of Burkholderia-associated diseases. This review provides a comprehensive and up-to-date account regarding the structural elucidation and biological activities of surface polysaccharides produced by Burkholderia species. The chemical synthesis of oligosaccharides mimicking Burkholderia polysaccharides is described in detail. Emphasis is placed on the recent research efforts toward the development of glycoconjugate vaccines against melioidosis and glanders based on synthetic or native Burkholderia oligo/polysaccharides.
Publication DOI: 10.1039/C8NP00046HJournal NLM ID: 8502408Publisher: London: Royal Society of Chemistry
Correspondence: charles.gauthier
iaf.inrs.ca
Institutions: INRS-Institut Armand-Frappier, Universite du Quebec, 531, boul. des Prairies, Laval, Canada
- Compound ID: 906
Structure type: homopolymer
Trivial name: 6-deoxy-heptan CPS
Compound class: CPS, O-polysaccharide, O-antigen
Reference(s) to other database(s): GTC:G09650IH, GlycomeDB:
3383
- Compound ID: 2243
Structure type: polymer chemical repeating unit
Trivial name: K-antigen, galactan-Kdo, Kdo-EPS
Compound class: CPS, EPS
Reference(s) to other database(s): GTC:G77110LE, GlycomeDB:
25312
- Compound ID: 3263
Structure type: homopolymer
Compound class: CPS, O-polysaccharide, mannan
Reference(s) to other database(s): GTC:G44405PH, GlycomeDB:
26469
- Compound ID: 12951
|
/Variants 1/-+ /Variants 0/-+
| |
a-L-6dTalp2Ac3Me4Ac-(1-3)-b-D-Glcp-(1-3)-{{{-a-L-6dTalp4(%)Ac-(1-3)-b-D-Glcp-(1-3)-}}}a-L-6dTalp4(%)Ac-(1-3)-b-D-Glcp
/Variants 0/ is:
?%Me-2)-
OR (exclusively)
?%Ac-2)-
/Variants 1/ is:
?%Me-2)-
OR (exclusively)
?%Ac-2)- |
Show graphically |
Structure type: oligomer
Compound class: O-antigen
- Compound ID: 12954
|
R-3HOMyr-(1-3)-+
|
?%b-L-Arap4N-(1--P--1)--+ |
| |
?%b-L-Arap4N-(1--P--4)--+ | |
| | |
Myr-(1-3)-R-3HOPam-(1-2)-b-D-GlcpN-(1-6)-a-D-GlcpN
| |
?%R-3HOMyr-(1-3)-+ |
|
R-3HOPam-(1-2)-+ |
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Structure type: oligomer
Compound class: lipid A
- Compound ID: 423
|
b-Kdop-(2-3)-+
|
-3)-b-D-Glcp-(1-4)-b-D-Glcp-(1-4)-a-L-6dTalp2Ac-(1- |
Show graphically |
Structure type: polymer chemical repeating unit
Compound class: CPS, EPS
Reference(s) to other database(s): GTC:G93694GD, GlycomeDB:
26572
- Compound ID: 549
Structure type: homopolymer
Trivial name: levan, levan-type polysaccharide
Compound class: EPS, polysaccharide, fructan
Reference(s) to other database(s): GTC:G29465PK, GlycomeDB:
9593
- Compound ID: 829
Structure type: polymer chemical repeating unit
Trivial name: common antigen, common polysaccharide antigen, A band polysaccharide, D-rhamnan, repeat unit of the lipopolysaccharide side-chain, A-band polysaccharide (A-PS), A-band polysaccharide
Compound class: O-polysaccharide, O-antigen, LPS, enterobacterial common antigen, CPA
Reference(s) to other database(s): GTC:G80135ZC, GlycomeDB:
25893
- Compound ID: 1294
Structure type: polymer chemical repeating unit
Compound class: O-polysaccharide, O-antigen
Reference(s) to other database(s): GTC:G40176QY, GlycomeDB:
25355
- Compound ID: 3724
Structure type: polymer chemical repeating unit
Trivial name: succinoglycan
Compound class: EPS
- Compound ID: 12955
|
b-D-Galp3(%)Ac4(%)Ac6(%)Ac-(1-6)-+
|
b-D-Galp2(%)Ac3(%)Ac4(%)Ac6(%)Ac-(1-2)-a-D-Rhap-(1-4)-+ |
| |
-3)-a-D-GlcpA-(1-3)-a-D-Manp-(1-3)-b-D-Glcp-(1-
|
a-D-Galp3(%)Ac4(%)Ac-(1-2)-+ |
Show graphically |
Structure type: polymer chemical repeating unit
Trivial name: cepacian
Compound class: EPS
Reference(s) to other database(s): GTC:G19685SV
- Compound ID: 12956
Structure type: polymer chemical repeating unit
Trivial name: co-cepacian
Compound class: EPS
Reference(s) to other database(s): GTC:G86277RB
- Compound ID: 12957
Structure type: polymer chemical repeating unit
Trivial name: PNAG
Compound class: EPS
Reference(s) to other database(s): GTC:G84392XC
- Compound ID: 2824
Structure type: polymer chemical repeating unit
Compound class: CPS, EPS
Reference(s) to other database(s): GTC:G84871IX, GlycomeDB:
25979
- Compound ID: 9210
|
-7)-a-Caryophyllosep5(%)Ac8(%)Ac9(%)Ac-(1-
Caryophyllose = 3,6,10-trideoxy-4-C-[(R)-1-hydroxyethyl]-D-erythro-D-gulo-decose |
Show graphically |
Structure type: homopolymer
Trivial name: caryophyllan
Compound class: O-polysaccharide, O-antigen
- Compound ID: 12967
|
-7)-b-Caryosep6(%)Ac-(1-
Caryose = 4,8-cyclo-3,9-dideoxy-L-erythro-D-ido-nonose |
Show graphically |
Structure type: homopolymer
Trivial name: caryan
Compound class: O-antigen
- Compound ID: 12968
|
L-gro-a-D-manHepp-(1-5)-a-Kdop-(2-4)-+
|
a-D-Glcp-(1-6)-b-D-Glcp-(1-4)-+ |
| |
L-gro-a-D-manHepp-(1-7)-+ | |
| | |
a-D-Galp-(1-2)-+ | | |
| | | |
Subst-(1-3)-b-QuipNAc-(1-6)-a-D-Glcp-(1-6)-a-D-Glcp-(1-3)-L-gro-a-D-manHepp-(1-3)-L-gro-a-D-manHepp-(1-5)-a-Kdop-(2--/(2->6) lipid A/
Subst = caryan (ID 12525) |
Show graphically |
Structure type: oligomer
Aglycon: (2->6) lipid A
Compound class: LOS
- Compound ID: 1295
Structure type: polymer chemical repeating unit
Compound class: O-polysaccharide, O-antigen
Reference(s) to other database(s): GTC:G43495GI, GlycomeDB:
25363
- Compound ID: 2796
|
-4)-a-D-Glcp-(1-2)-a-L-Rhap-(1-4)-a-D-GlcpA-(1-3)-b-L-Rhap2Ac-(1-4)-b-D-Glcp-(1- |
Show graphically |
Structure type: polymer chemical repeating unit
Compound class: EPS
Reference(s) to other database(s): GTC:G70458LL, GlycomeDB:
25969
- Compound ID: 8947
|
b-D-Galp-(1-6)-+
|
b-D-Glcp-(1-3)-+ |
| |
b-D-Galp-(1-2)-a-D-Rhap-(1-4)-+ |
| |
-3)-a-D-GlcpA-(1-3)-a-D-Manp-(1-3)-b-D-Glcp-(1-
|
a-D-Galp-(1-2)-+ |
Show graphically |
Structure type: polymer chemical repeating unit
Compound class: EPS
Reference(s) to other database(s): GTC:G98289QM, GlycomeDB:
37864
- Compound ID: 2040
Structure type: polymer chemical repeating unit
Compound class: O-polysaccharide, O-antigen
Reference(s) to other database(s): GTC:G03826MN, GlycomeDB:
25174
- Compound ID: 1279
Structure type: polymer chemical repeating unit
Compound class: O-polysaccharide, O-antigen
Reference(s) to other database(s): GTC:G89062YH, GlycomeDB:
3437
- Compound ID: 1280
Structure type: polymer chemical repeating unit
Compound class: O-polysaccharide, O-antigen
Reference(s) to other database(s): GTC:G78324BK, GlycomeDB:
2534
- Compound ID: 2041
Structure type: polymer chemical repeating unit
Compound class: O-polysaccharide, O-antigen, LPS
Reference(s) to other database(s): GTC:G91041IQ, GlycomeDB:
25175
- Compound ID: 2111
|
a-L-Rhap-(1-2)-+
|
-3)-a-D-Fucp-(1-3)-a-D-Fucp-(1-3)-a-L-Rhap-(1- |
Show graphically |
Structure type: polymer chemical repeating unit
Compound class: O-polysaccharide, O-antigen
Reference(s) to other database(s): GTC:G29596PG, GlycomeDB:
25232
- Compound ID: 12959
|
a-L-Rhap-(1-2)-+ b-D-Glcp-(1-4)-+ a-Kop-(2-4)-+
| | |
L-gro-a-D-manHepp-(1-7)-L-gro-a-D-manHepp-(1-3)-L-gro-a-D-manHepp-(1-5)-a-Kdop |
Show graphically |
Structure type: oligomer
Compound class: core oligosaccharide
Reference(s) to other database(s): GTC:G60958CB
- Compound ID: 940
Structure type: polymer chemical repeating unit
Compound class: O-polysaccharide, O-antigen
Reference(s) to other database(s): GTC:G50210GF, GlycomeDB:
27103
- Compound ID: 3455
Structure type: polymer chemical repeating unit
Trivial name: 6-deoxy-D-talan
Compound class: EPS
Reference(s) to other database(s): GTC:G37166XU, GlycomeDB:
26840
- Compound ID: 324
Structure type: polymer chemical repeating unit
Compound class: O-polysaccharide, O-antigen
Reference(s) to other database(s): GTC:G42886IQ, GlycomeDB:
25555
- Compound ID: 6947
Structure type: polymer chemical repeating unit
Compound class: O-polysaccharide, O-antigen, polysaccharide
Reference(s) to other database(s): GTC:G22669IM, GlycomeDB:
27888
- Compound ID: 6948
Structure type: polymer chemical repeating unit
Compound class: EPS, polysaccharide
Reference(s) to other database(s): GTC:G98789SQ, GlycomeDB:
27889
- Compound ID: 10649
|
L-gro-a-D-manHepp-(1-6)-b-D-Glcp-(1-4)-+
|
b-D-QuipNAc-(1-3)-a-L-Rhap-(1-2)-+ | a-Kop-(2-4)-+
| | |
b-D-Glcp-(1-3)-a-D-GalpNAc-(1-3)-b-D-GalpNAc-(1-3)-a-L-Rhap-(1-3)-L-gro-a-D-manHepp-(1-3)-L-gro-a-D-manHepp-(1-5)-a-Kdop
|
L-gro-a-D-manHepp-(1-7)-+ |
Show graphically |
Structure type: oligomer
Compound class: core oligosaccharide
Reference(s) to other database(s): GTC:G03152EV
- Compound ID: 8194
|
?%b-D-Xylp-(1-2)-+
|
-3)-a-L-6dTalp-(1-3)-b-D-GalpNAc-(1-
|
?%b-D-Xylp-(1-4)-+ |
Show graphically |
Structure type: polymer chemical repeating unit
Compound class: O-polysaccharide, O-antigen
Reference(s) to other database(s): GTC:G40227PF, GlycomeDB:
37114
- Compound ID: 8430
Structure type: polymer chemical repeating unit
Compound class: O-polysaccharide, O-antigen
Reference(s) to other database(s): GTC:G61497FY, GlycomeDB:
37265
- Compound ID: 12966
|
L-gro-a-D-manHepp-(1-7)-L-gro-a-D-manHepp-(1-7)-+ b-D-Glcp-(1-4)-+ a-Kop-(2-4)-+
| | |
b-D-Galp-(1-3)-a-D-GalpNAc-(1-3)-b-D-GalpNAc-(1-3)-L-gro-a-D-manHepp-(1-3)-L-gro-a-D-manHepp-(1-5)-a-Kdop-(2--/(2->6) lipid A/
|
a-L-Rhap-(1-2)-+ |
Show graphically |
Structure type: oligomer
Aglycon: (2->6) lipid A
Compound class: LOS
Reference(s) to other database(s): GTC:G08282ML
- Compound ID: 10589
Structure type: polymer chemical repeating unit
Compound class: O-polysaccharide, O-antigen
Reference(s) to other database(s): GTC:G12239HW
- Compound ID: 12186
|
-3)-a-D-Manp-(1-2)-a-D-Manp-(1-2)-a-D-Rhap3(50%)Me-(1-3)-a-D-Rhap-(1- |
Show graphically |
Structure type: polymer chemical repeating unit
Trivial name: rhamno-mannan, rhamnomannan, Epol C1576
Compound class: EPS
Reference(s) to other database(s): GTC:G72472ZR
- Compound ID: 12961
|
L-Arap4N-(1-8)-a-Kop-(2-4)-+
|
L-gro-a-D-manHepp-(1-7)-+ b-D-Glcp-(1-4)-+ |
| | |
b-D-QuipNAc-(1-7)-L-gro-a-D-manHepp-(1-2)-a-D-Glcp-(1-3)-L-gro-a-D-manHepp-(1-3)-L-gro-a-D-manHepp-(1-5)-a-Kdop-(2--/(2->6) lipid A/ |
Show graphically |
Structure type: oligomer
Aglycon: (2->6) lipid A
Compound class: LOS
Reference(s) to other database(s): GTC:G54799XB
- Compound ID: 240
Structure type: polymer chemical repeating unit
Trivial name: neutral glycan
Compound class: O-polysaccharide, O-antigen, LPS
Reference(s) to other database(s): GTC:G46905WE, GlycomeDB:
3417, CCSD:
12532, CBank-STR:2856
- Compound ID: 12965
|
b-L-Arap4N-(1-8)-a-Kop-(2-4)-+
|
a-L-Rhap-(1-2)-+ b-D-Glcp-(1-4)-+ |
| | |
D-gro-a-D-manHepp-(1-7)-L-gro-a-D-manHepp-(1-3)-L-gro-a-D-manHepp-(1-7)-L-gro-a-D-manHepp-(1-3)-L-gro-a-D-manHepp-(1-5)-a-Kdop-(2--/(2->6) lipid A/ |
Show graphically |
Structure type: oligomer
Aglycon: (2->6) lipid A
Compound class: LOS
Reference(s) to other database(s): GTC:G04017UD
- Compound ID: 12963
|
a-D-Galp-(1-6)-b-D-Glcp-(1-4)-+
|
L-gro-a-D-manHepp-(1-7)-+ |
| |
a-D-Glcp-(1-6)-a-D-Glcp-(1-3)-L-gro-a-D-manHepp-(1-3)-L-gro-a-D-manHepp-(1-5)-a-Kdop-(2--/(2->6) lipid A/ |
Show graphically |
Structure type: oligomer
Aglycon: (2->6) lipid A
Compound class: LOS
Reference(s) to other database(s): GTC:G84208RC
- Compound ID: 12952
|
/Variants 1/-+ /Variants 0/-+
| |
a-L-6dTalp2Ac3Me-(1-3)-b-D-Glcp-(1-3)-{{{-a-L-6dTalp-(1-3)-b-D-Glcp-(1-3)-}}}a-L-6dTalp-(1-3)-b-D-Glcp
/Variants 0/ is:
?%Me-2)-
OR (exclusively)
?%Ac-2)-
/Variants 1/ is:
?%Me-2)-
OR (exclusively)
?%Ac-2)- |
Show graphically |
Structure type: oligomer
Compound class: O-antigen
- Compound ID: 10818
Structure type: polymer chemical repeating unit
Compound class: O-polysaccharide, O-antigen
Reference(s) to other database(s): GTC:G46744WV
- Compound ID: 10819
Structure type: homopolymer
Compound class: O-polysaccharide, O-antigen
Reference(s) to other database(s): GTC:G09327TR
- Compound ID: 11118
|
a-D-Glcp-(1-4)-a-D-GlcpA-(1-3)-a-L-Fucp-(1-3)-+
|
-3)-b-D-Glcp-(1-4)-a-D-Glcp-(1-3)-a-L-Fucp-(1- |
Show graphically |
Structure type: polymer chemical repeating unit
Compound class: EPS
Reference(s) to other database(s): GTC:G73833FX
- Compound ID: 12964
|
L-gro-a-D-manHepp-(1-7)-+
|
b-D-Galp-(1-3)-a-D-Glcp-(1-6)-+ |
| |
b-D-QuipNAc-(1-7)-+ | | b-D-Glcp-(1-4)-+ a-Kop-(2-4)-+
| | | | |
L-gro-a-D-manHepp-(1-3)-L-gro-a-D-manHepp-(1-2)-a-D-Galp-(1-3)-L-gro-a-D-manHepp-(1-3)-L-gro-a-D-manHepp-(1-5)-a-Kdop-(2--/(2->6) lipid A/
|
b-D-Glcp-(1-2)-+ |
Show graphically |
Structure type: oligomer
Aglycon: (2->6) lipid A
Compound class: LOS
Reference(s) to other database(s): GTC:G24961HK
- Compound ID: 4664
Structure type: homopolymer
Compound class: O-polysaccharide, O-antigen
Reference(s) to other database(s): GTC:G63740AV, GlycomeDB:
2516
- Compound ID: 12971
|
a-D-Galp-(1-3)-+
|
-3)-a-D-Manp2(%)Ac-(1-3)-b-D-Glcp-(1-4)-a-D-Manp-(1- |
Show graphically |
Structure type: polymer chemical repeating unit
Compound class: EPS
Reference(s) to other database(s): GTC:G31490UJ
- Compound ID: 12973
|
L-gro-a-D-manHepp-(1-6)-b-D-Glcp-(1-4)-+
|
L-gro-a-D-manHepp-(1-7)-+ | a-Kop-(2-4)-+
| | |
b-D-GalpNAc-(1-3)-a-L-Rhap-(1-3)-L-gro-a-D-manHepp-(1-3)-L-gro-a-D-manHepp-(1-5)-a-Kdop-(2--/(2->6) lipid A/ |
Show graphically |
Structure type: oligomer
Aglycon: (2->6) lipid A
Compound class: LOS
Reference(s) to other database(s): GTC:G27801RM
- Compound ID: 12456
|
b-L-Xylp-(1-2)-+
|
-2)-a-L-Rhap-(1-3)-b-L-Rhap-(1-4)-a-L-Rhap-(1- |
Show graphically |
Structure type: polymer chemical repeating unit
Compound class: O-polysaccharide, O-antigen
Reference(s) to other database(s): GTC:G58661KN
- Compound ID: 12953
|
a-L-6dTalp2Ac3Me4Ac-(1-3)-b-D-Glcp-(1-3)-{{{-a-L-6dTalp2Ac-(1-3)-b-D-Glcp-(1-3)-}}}a-L-6dTalp2Ac-(1-3)-b-D-Glcp |
Show graphically |
Structure type: oligomer
Compound class: O-antigen
Reference(s) to other database(s): GTC:G64097ZD
- Compound ID: 7098
Structure type: polymer chemical repeating unit
Compound class: O-polysaccharide, O-antigen
Reference(s) to other database(s): GTC:G87187IQ, GlycomeDB:
27848
- Compound ID: 1283
Structure type: polymer chemical repeating unit
Compound class: O-polysaccharide, O-antigen
Reference(s) to other database(s): GTC:G11401HK, GlycomeDB:
3435, CCSD:
12565, CBank-STR:2460
- Compound ID: 1277
Structure type: polymer chemical repeating unit
Compound class: O-polysaccharide, O-antigen
Reference(s) to other database(s): GTC:G84739NX, GlycomeDB:
3434
- Compound ID: 26
Structure type: polymer chemical repeating unit
Compound class: O-polysaccharide, O-antigen
Reference(s) to other database(s): GTC:G37099RV, GlycomeDB:
25434
- Compound ID: 27
Structure type: polymer chemical repeating unit
Trivial name: mannan
Compound class: O-polysaccharide, O-antigen
Reference(s) to other database(s): GTC:G40440IC, GlycomeDB:
27903
- Compound ID: 4900
Structure type: polymer chemical repeating unit
Compound class: O-polysaccharide, O-antigen
Reference(s) to other database(s): GTC:G94962EH, GlycomeDB:
3440
- Compound ID: 1282
Structure type: polymer chemical repeating unit
Compound class: O-polysaccharide, O-antigen
Reference(s) to other database(s): GTC:G62219GM, GlycomeDB:
3439
- Compound ID: 1293
Structure type: polymer chemical repeating unit
Compound class: O-polysaccharide, O-antigen
Reference(s) to other database(s): GTC:G78814IK, GlycomeDB:
3436
- Compound ID: 1291
Structure type: polymer chemical repeating unit
Compound class: O-polysaccharide, O-antigen
Reference(s) to other database(s): GTC:G86024VK, GlycomeDB:
3447
- Compound ID: 1290
Structure type: polymer chemical repeating unit
Compound class: O-polysaccharide, O-antigen
Reference(s) to other database(s): GTC:G35276WU, GlycomeDB:
3446
- Compound ID: 12958
Structure type: polymer chemical repeating unit
Compound class: O-antigen
Reference(s) to other database(s): GTC:G19567BD
- Compound ID: 1292
Structure type: polymer chemical repeating unit
Compound class: O-polysaccharide, O-antigen
Reference(s) to other database(s): GTC:G96363IL, GlycomeDB:
3438
- Compound ID: 1284
Structure type: polymer chemical repeating unit
Compound class: O-polysaccharide, O-antigen
Reference(s) to other database(s): GTC:G99774AJ, GlycomeDB:
3441
- Compound ID: 1285
Structure type: polymer chemical repeating unit
Compound class: O-polysaccharide, O-antigen
Reference(s) to other database(s): GTC:G51061VL, GlycomeDB:
3442
- Compound ID: 1286
Structure type: polymer chemical repeating unit
Compound class: O-polysaccharide, O-antigen
Reference(s) to other database(s): GTC:G04976QE, GlycomeDB:
3443
- Compound ID: 1287
Structure type: polymer chemical repeating unit
Trivial name: b-specific polysaccharide antigen
Compound class: O-polysaccharide, O-antigen, S-layer glycoprotein
Reference(s) to other database(s): GTC:G48878CJ, GlycomeDB:
3444
- Compound ID: 11153
Structure type: polymer chemical repeating unit
Compound class: O-polysaccharide, O-antigen
Reference(s) to other database(s): GTC:G22212ZI
- Compound ID: 11154
Structure type: polymer chemical repeating unit
Compound class: O-polysaccharide, O-antigen
Reference(s) to other database(s): GTC:G83770MG
- Compound ID: 12960
|
L-gro-a-D-manHepp-(1-5)-a-Kop-(2-4)-+
|
b-D-Glcp-(1-4)-+ |
| |
EtN-(1--P--4)--a-D-Glcp-(1-3)-L-gro-a-D-manHepp-(1-3)-D-gro-a-D-manHepp-(1-5)-a-Kdop-(2--/(2->6) lipid A/ |
Show graphically |
Structure type: oligomer
Aglycon: (2->6) lipid A
Compound class: LOS
Reference(s) to other database(s): GTC:G11325BQ
- Compound ID: 12969
|
-2)-a-L-Rhap-(1-2)-a-L-Rhap3Me-(1-2)-a-L-Rhap-(1-4)-b-D-Galp2Me-(1- |
Show graphically |
Structure type: polymer chemical repeating unit
Compound class: O-antigen
Reference(s) to other database(s): GTC:G55559IL
- Compound ID: 12970
|
-2)-a-L-Rhap-(1-2)-a-L-Rhap3Me-(1-2)-a-L-Rhap-(1-4)-b-D-Galp-(1- |
Show graphically |
Structure type: polymer chemical repeating unit
Compound class: O-antigen
Reference(s) to other database(s): GTC:G16446RA
- Compound ID: 12974
|
a-D-GlcpA-(1-4)-+
|
-4)-b-D-Glcp-(1-2)-a-L-Rhap-(1-4)-a-D-Glcp-(1-3)-b-L-Rhap2Ac-(1- |
Show graphically |
Structure type: polymer chemical repeating unit
Compound class: EPS, O-polysaccharide, glucuronorhamnoglucan
Reference(s) to other database(s): GTC:G26601LQ
Found